Similarly, indirectly regulated genes, ATF5 and CHD4 that participate in maintaining centrosome integrity (Sillibourne et al., 2007; Madarampalli et al., 2015), could also have additional ciliary roles as transcription factors in cilia [defects in centrosome structure or function may lead to ciliopathies; Bettencourt-Dias et al., 2011)]. Here, ATF5 is linked to ciliopathy.