CTSB and Parkinson disease: Most of these were located in loci with suggestive associations in PD GWAS (5 × 10−8 < p < 1 × 10−6, Supplementary Figures 11–49), and four genes (CTSB, PDLIM2, GALC, and C8orf5) were previously found to be genome-wide significant in PD GWAS meta-analysis3.