Contrast colocalization analysis for the meconium ileus GWAS loci showed that eQTLs for SLC6A14, SLC26A9 and ATP12A in the pancreas colocalize with meconium ileus-associated variants significantly more than eQTLs for these genes in the other meconium ileus-relevant tissues tested including the intestine (Table 3, SSC colocalization p-value results). Here, SLC26A9 is linked to intestinal obstruction in the newborn due to guanylate cyclase 2C deficiency.