DBA staining of the cyst wall is principally due to O-glycosylation of the mucin-like glycoprotein CST1, since deletion of CST1, pp-GalNAcT2, or pp-GalNAcT3 led to a loss of staining by this lectin (Tomita et al., 2013, 2017). The gene discussed is CST1; the disease is cyst.