PPY and cyst: Although PpY-Gal4 is expressed in the SCCs from the post-mitotic stages (Jung et al., 2007), PpY-Gal4-mediated expression of dsDlg1 eliminates the protein from the SCCs only at the last stage of spermatid maturation, when the cyst enters the TE, and leads to premature spermatid release inside the testis.