DCLK1 and helminthiasis: In addition, expression of the epithelial tuft cell markers DCLK1 and SOX9 were increased in T. suis-infected pigs fed dietary inulin, highlighting the possible role of inulin in Th2 polarization, as tuft cells have recently been shown to be crucial players in initiation of Th2 responses during helminth infection in mice (34).