GNL3 and bipolar disorder: In addition, a transcript annotated to GNL3, encoding G Protein Nucleolar 3, was associated with both bipolar disorder (P-SMR = 2.17 × 10− 6) and schizophrenia (P-SMR = 2.71 × 10− 7), with risk alleles for both disorders associated with increased expression of this transcript in fetal brain.