CLIP1 and infection: Related to the movements of capsids within infected cells, other reports suggest that HSV-1 engages dynamic microtubules (MTs) at early stages of infection, using plus end-tracking proteins (+TIPs) complexes composed of cytoplasmic linker protein 170 (CLIP-170), dynactin-1 (DCTN1) and end-binding protein (EB1), with retrograde capsid transport being dependent on CLIP-170 (Jovasevic et al., 2015).