For example, citrullinated H3 is present on LDs in early mouse oocytes (Kan et al., 2012); and H2A and H2B, were identified as high confidence LD proteins in human osteosarcoma (U2OS) and human hepatocellular carcinoma (Huh7) cell lines (Bersuker et al., 2018). Here, H2BC21 is linked to hepatocellular carcinoma.