Analysis of tracheal organ rings at 4 days post-infection from the chickens infected with the rIBVs based on Beaudette but with modified ADRP domains, BeauR-G-ADRP, BeauR-M41-ADRP, BeauR-M41(S)-G-ADRP and BeauR-M41(S)-M41-ADRP, showed average (mean) ciliary activities of 84, 74, 78 and 71 %, respectively, comparable to those from the groups infected with Beau-R (76 %) and BeauR-M41(S) (78 %). Here, PLIN2 is linked to infection.