Quantification of expression on immunolabelled tissue sections with whole slide imaging and image analysis [6] corroborates the RNA expression data of Pten/p53 and Rb/p53ant tumours with their high expression of Olig2, PDGFRα and Sox2, whilst Rb/p53 grafts expressed much lower levels of these markers with quantification data in the right column (Figs. 6g, k, o, s). This evidence concerns the gene OLIG2 and neoplasm.