TUSC2P (tumor suppressor 2, mitochondrial calcium regulator pseudogene) regulated TUSC2 (tumor suppressor 2, mitochondrial calcium regulator), as well as TIMP2 and TIMP3 (tissue inhibitor of metalloproteinases 2 and 3) by sequestering multiple miRNAs, including miR-17, miR-93, miR-299-3p, miR-520a, miR-608 and miR-661, to reduce the growth and migratory capacity of various cancer cell lines [128]. Here, TIMP2 is linked to cancer.