GREM1 and infection: Interestingly, following infection with the EM control virus, T-MSCs displayed increased expression only of GREM1 and ITGA11 compared to N-MSCEM, thereby displaying a slightly different profile than non-infected T-MSCs (Fig. 3B and S7B) and possibly explaining the smaller difference in metastasis promotion between T- and N-MSCEM than between the corresponding uninfected MSCs.