Solorzano et al. (2015) have indeed demonstrated that Grp is upregulated in DRG neurons upon nerve injury (Solorzano et al., 2015). If nerves were injured during the first injection, TVA may have been expressed in DRG neurons, including neurons of the NP subtype. These neurons would then have been susceptible to direct infection by the EnvA-pseudotyped rabies. Finally, Sun et al. (2017) have used a 7 d incubation time, which was 2 d more than the incubation time we chose for this study. Here, GRP is linked to infection.