UGP2 and congenital secretory chloride diarrhea 1: Of these seven candidate genes, three of the genes/loci were identified from both the GH2010 and Field2011 environments: r locus, UGPase, and AGPS2. The polymorphisms in the seven genes/loci were associated with CLD at DP10, DP16, DP17, DP34, and DP39 in the GH2010 environment; and with DP17, DP18 and DP23 in the Field2011 environment.