BMP3 and Brachycephaly: Elsewhere, measurements and geometric morphometrics were used to quantify skull shape, revealing quantitative trait loci (QTL) associated with brachycephaly on CFA1, CFA5, CFA18, CFA30, and CFAX and a missense variant in the bone morphogenetic protein 3 (BMP3) gene on CFA32 [8, 9].