Among the 24 loci associated with triglyceride-related traits in the present study, 13 loci—BUD13 [15], 11q23.3 [16], APOA5 [15, 16], ZPR1 [16], APOA4 [15, 17], LPL [15, 17], 8p21.3 [15, 18], SIK3 [19, 20], GCKR [16], 2p23 [17, 21], C2orf16 [22], 8q24.1 [23], and LOC101929011 [21]—were previously shown to be related to the circulating triglyceride level or hypertriglyceridemia (Supplementary Table 16). Here, APOA4 is linked to hypertriglyceridemia.