ART4 and cancer: We have previously shown in two separate studies, respectively, that ovarian cancer cells, IGROV1 and SKOV3, have higher levels of glycoprotein α2-6 sialylation as compared to non-cancer ovarian epithelial cells19 and nLc4 and P1 are both expressed on IGROV1 and ovarian cancer tissue-derived GSL-glycans13.