OLR1 and benign prostatic hyperplasia: Although the four major BPH resistance QTLs were cloned by map-based techniques, many BPH resistance-related genes, including OsHI-LOX [12], Bphi008a [13], OsERF3 [14], OsPLD α4 and α5 [15], OsACS2 [17], Osr9-LOX1 [19], and OsJMT1 [20], were also isolated using a reverse genetics strategy.