Similarly, 7 of the 9 SNPs significantly heterogeneous between T1R and leprosy were eQTLs in either whole blood, rs3774937 (NFKB1), rs10065637 (ANKRD55), rs11150589 (ITGAL) and rs2836878 (lncRNA ENSG00000235888) or multiple tissues, rs4664304 (LY75), rs113653754 (HLA-DQB1) and rs4768236 (LRRK2; S4 Fig) [19, 28]. Here, LRRK2 is linked to leprosy.