According to the Geneimprint database (http://www.geneimprint.com/site/genes-byspecies), five imprinted genes are mapped to 7q32: COPG2IT1, MEST, CPA4, MESTIT1 and KLF14, of which, a loss of imprinting (biallelic expression) of MEST has been associated with BC development40, 41, 42. Here, MEST is linked to breast cancer.