ERAP1 and Behcet disease: Notably, among the genetic loci implicated in previous GWAS2, 3, 4, 5, 6, 16, IL10 and ERAP1 also showed modest associations with intestinal BD (rs1518111, combined P = 7.6 × 10−3, Table 3; rs2927615, combined P = 1.6 × 10−2), and HLA-B (near PSORS1C1) was significantly associated with BD without intestinal involvement compared to healthy controls (rs4959053, combined P = 6.6 × 10−5, P replication = 4.6 × 10−3; rs12525170, combined P = 1.3 × 10−5, P replication = 1.1 × 10−5, Table 4).