The aggregates were observed with non-hydroxamic acid HDAC inhibitors, such as CXD101, and occurred in other cell lines, including the human DLBCL cell lines RIVA and HBL-1 and the neuroblastoma SH-SY5Y cell line (Figures 2, 4, S1, S2, and S5). Here, HDAC9 is linked to neuroblastoma.