BBS9 and leprosy: Through the meta-analysis of a total number of 8,156 leprosy patients and 15,610 healthy controls, we have identified four novel associations, all of which can indicate candidate genes within the susceptibility loci, SYN2 on 3p25.2, BBS9 on 7p14.3, CTSB on 8p23.1 and MED30 on 8q24.11, through a differential gene expression and eQTL analysis.