PSMB9 and Epstein-Barr virus infection: When we performed the neutralization assay in HEK-293 epithelial cells, sera from mice immunized with UV-inactivated EBV at the 1:2.5 dilution effectively neutralized 94.3% of EBV infection, followed by sera from mice immunized with gH/gL-EBNA1 VLPs (77.9%), gB-LMP2 VLPs (76.8%), and gp350/220 VLPs (54.9%) (Figure 5C).