Graphical genotyping was also successful in showing the haplotypes on chromosome 12 carrying Ry-fsto, another resistance gene derived from S. stoloniferum conferring broad-spectrum resistance to PVY, as well as chromosome 5 haplotypes from S. vernei, with the Gpa5 locus involved in resistance against Globodera pallida cyst nematodes. The gene discussed is HSPA4; the disease is cyst.