In addition, the genes prioritized for RA included 5/9 candidates (C8Orf13, BLK, TRAF1, FADS2, and SYNGR1) that were identified in a recent study (Zhu et al., 2016) combining whole-blood eQTL with RA GWAS data by Mendelian randomization. Here, SYNGR1 is linked to rheumatoid arthritis.