SSX2IP and cancer: Of note, abnormal accumulation of satellite particles in the absence of microtubule anchoring (i.e. MSD1/SSX2IP depletion) is observed only in cancer-derived culture cells (U2OS, HeLa, MCF-7, A549, T98G and Saos-2); non-transformed cells (RPE1, W138 and MG00024B) do not display satellite aggregation [28].