LGALS4 and cyst: In addition, 73 (65%) of the 113 lines had abnormal GSC phenotypes with Nos-Gal4 and c587-Gal4, similar to their phenotypes with Act-Gal4, suggesting that 65% of the genes regulate GSC fates intrinsically or externally from CySCs or local early cyst cells, and the other 35% regulate GSCs from distant organs.