Moreover, doxocyclin-induced overexpression of ARTD9 in an ARTD9-negative low-risk GCB-DLBCL cell line led to the induction of a small subset of IFN(γ)-related genes [469], thus provided preliminary evidence that ARTD9 might play a direct role in IFN signaling pathways [469]. Here, PARP9 is linked to diffuse large B-cell lymphoma.