Cathepsin L (CTSL), which primes the filovirus glycoprotein in the endosome [22], EXT1 [23], which is involved in biosynthesis of heparan sulfate, and Niemann-Pick disease, type C1 (NPC1 [24, 25]), showed strong bias against MARV entry (Table 1). Here, ART4 is linked to frontotemporal dementia.