TARDBP and frontotemporal dementia: However, in ALS/FTD human samples, there are a handful of examples of co-labeling of TDP-43 and FUS inclusions with well-known SG markers (Colombrita et al., 2009; Volkening et al., 2009; Dormann et al., 2010; Liu-Yesucevitz et al., 2010; Bentmann et al., 2012).