For example, in addition to NNAT and PSMB5, eQTLs for a long non-coding RNA (PKI55; rs13392737), adaptor related protein complex 1, sigma 1 subunit (AP1S1; rs10279545) and translocation associated membrane protein 1 (TRAM1; rs2959574) were also associated with AD at p = p = 9.1x10-4 and p = 6.2x10-3, respectively. Here, PSMB5 is linked to Alzheimer disease.