Together these data demonstrate a critical role for groups of ciliopathy genes including AHI1, ALMS1, BBS1, BBS4, BBS7, BBS9, BBS10, BBS11 [TRIM32], BBS12, BUBR1 [BUB1B], IFT80, KIF7, NPHP1, NPHP8 [RPGRIP1L], TCTN2, TMEM216 and TUB in the coordination of migration and placement of neurons in the developing cerebral cortex. Here, BBS12 is linked to ciliopathy.