Consistent with the migration defects, we found an aberrant laminar placement of neurons following disruption of 17 ciliopathy genes (AHI1, ALMS1, BBS1, BBS4, BBS7, BBS9, BBS10, BBS11 [TRIM32], BBS12, BUBR1 [BUB1B], IFT80, KIF7, NPHP1, NPHP8 [RPGRIP1L], TCTN2, TMEM216 and TUB). Here, BBS12 is linked to ciliopathy.