MUC5AC and inflammatory bowel disease: Alterations in mucin glycosylation have been associated with a number of diseases such as colitis, colonic cancer and inflammatory bowel diseases in humans (e.g., McGovern et al., 2010; Larsson et al., 2011; Rausch et al., 2011; Parmar et al., 2012; Forni et al., 2014) and mouse models (e.g., An et al., 2007; Stone et al., 2009; Fu et al., 2011).