GRHL3 and Alzheimer disease: For instance, decrease in PV-expressing neurons occurs later and is less pronounced in the course of AD-like pathology than the loss of other types of GABAergic neurons (e.g., SOM- or calretinin-expressing neurons) in both olfactory bulb (Saiz-Sanchez et al., 2013) and olfactory cortex (Saiz-Sanchez et al., 2012) of the APPxPS1 mice.