Specifically, five acyl-CoA dehydrogenases (RC1_0378, RC1_1632, RC1_2142, RC1_3098, RC1_3534), two acetyl CoA acetyltransferases (RC1_0397, RC1_3948), two fatty acid desaturases (RC1_0393, RC1_1736), and an inositol-1-phosphate synthase (RC1_0768) are all up regulated 4- to 8-fold late in cyst development (Figure 4). Here, CBX8 is linked to cyst.