They noted that 4/61 affected dogs did not have the mutation and 9/38 normal controls did (attributed to low penetrance), and recently, Meurs and others (2014) have also reported that 7/43 genotyped Boxers diagnosed with ARVC were negative for the STRN mutation (16 per cent). Here, STRN is linked to Arrhythmogenic right ventricular dysplasia.