4 Kono Kono JO JO Podskarbi Podskarbi T T Shin Shin Y Y Lanzl Lanzl I I Oligosymptomatic cornea verticillata in a heterozygote for Fabry disease: a novel mutation in the alpha-galactosidase gene Oligosymptomatic cornea verticillata in a heterozygote for Fabry disease: a novel mutation in the alpha-galactosidase gene Cornea Cornea 2003 2003 22 22 2 2 175 175 177 177 12605057 12605057. Here, GLA is linked to Fabry disease.