In addition, the nuclear protein levels of MEOX2, HDAC9 and TWIST1 were consistently increased in the LT-derived samples (Figure 2B) as well as MEOX2 and TWIST1 protein levels/abundance in NSCLC cell lines (Figure S4A-C), whereas the expression of these proteins in the adjacent pulmonary parenchyma and the LNAT-derived samples were not increased (Figure 2B), as detected using immunohistochemistry. Here, TWIST1 is linked to non-small cell lung carcinoma.