In addition, the MSA-oligodendroglia containing a GCI (i.e., a pα-syn-positive cytoplasmic structure) had much less TPPP in the nucleus (types 3 + 5/types 3 + 4 + 5 + 6 in MSA = 19.6 ± 10.9%) than in the GCI-negative MSA-oligodendroglia (type 1/types 1 + 2 in MSA = 48.63 ± 10.37%, p < 0.001). Here, TPPP is linked to multiple system atrophy.