To assess further the role of Meis1 in skin carcinogenesis, we examined the expression of Meis1 in chemically induced papillomas, carcinomas in situ, and metastatic carcinomas from wild-type FVB/N mice by semi-quantitative RT-PCR and RNA sequencing (Aoto et al., unpublished data). Here, MEIS1 is linked to in situ carcinoma.