In contrast to Meis1 expression in normal epidermis (Fig. 1), we found that Meis1-EGFP was not co-expressed with β4 integrin, CD34, or K15 in the tumor basal layers, where epidermal cancer stem cells are thought to reside in papillomas (Fig. 7G–7O). This evidence concerns the gene MEIS1 and neoplasm.