CDK1 and hepatocellular carcinoma: The USF1 element in Cdc2 described in A549 epithelial cell line derived from a lung carcinoma tissue-, H1-hESC embryonic stem cells and HepG2 hepatocellular carcinoma was in the region +1044 to +1300 from the TSS, differently positioned from what we found for USF1 recruitment (Fig. 5D).