BSG and embryonal carcinoma: Miyauchi et al. discovered that Lotus tetragonolobus agglutinin (LTA) bound to CD147 from embryonal carcinoma cells and Kato et al. found that CD147 served as a ligand for E-selectin which recognizes sialylated glycans, such as Lewis X (sLex), both implying the sialyl Lewis X structure, namely, Galβ1, 4Fucα1,3GlcNAc, in N-glycans of CD147 [27,29].