MMP9 and gastric carcinoma: 30 showed that invasive behavior of GC cells was inhibited by treatment with anti-α6 integrin antibody. Saito et al. 31 reported that the production of MMP-9 by MKN1 human gastric carcinoma cells was potentiated by the α3β1 integrin–LN-5 interaction, which facilitated their invasion through degradation of the matrix.