HLA-DRB1 and sarcoidosis: In that study, subsequent conditional analyses revealed four additional independent variants (SNPs rs146146117 HLA-DQA1, rs9461776 HLA-DRB1, rs715299 NOTCH4, and rs9272320 HLA-DQA1) associated with sarcoidosis risk in the HLA class II region at the suggestive GWA significance threshold.