In our GWAS, multiple genetically-identical (r2 = 1) SNPs were significantly associated with sarcoidosis risk [14], including a missense SNP (rs7192) in HLA-DRA that has been associated with risk of both rheumatoid arthritis and systemic lupus erythematosus [26], and a splice-acceptor variant (rs8084) associated with rheumatoid arthritis [27], [28]. Here, HLA-DRA is linked to rheumatoid arthritis.