GATA4 and hepatocellular carcinoma: However, no significant difference was found with respect to their expression between normal hepatocytes and HCC cell lines (mRNA/protein: GATA4, p = 0.667/0.558; GATA5, p = 0.111/0.438; GATA6, p = 0.333/0.143; respectively, Fig. 1A and D).