BLMH and neoplasm: We tested the subset of the CGH samples (7 greyhounds and 7 Rottweilers) also included in the GWAS analysis and found that five of the top 29 GWAS SNPs are moderately associated with tumor gain or loss, most significantly at the gene BLMH, a candidate tumor suppressor gene for hepatocellular carcinoma [58] (Additional file 1: Table S5).