STEAP4 and obesity due to melanocortin 4 receptor deficiency: The STEAP4 gene encoding metalloreductase, which is associated with obesity and insulin-resistance in human [64]–[66], is located at 74.98 Mb on SSC9, very close to the strongest association signal (ALGA0053636) detected in the Sutai piglets, and thereby is an excellent positional and biological candidate gene for this QTL.