LGALS4 and autism: However, Gregory et al. [19] have indicated that CpG-934, whose differential methylation has also been associated with autism, falls within predicted binding domains for v-rel reticuloendotheliosis viral oncogene homolog (avian) (c-Rel), zinc fingers and homeoboxes 2 (ZHX2), and lectin, galactoside-binding, soluble, 4 (LGALS4).